Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China; College of Veterinary Medicine, Gansu Agricultural University, 730030, Lanzhou, China; Department of Experiment & Teaching, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: xbmzyd@163.com.

Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: 2864746700@qq.com.

Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: jiaminwang1987@163.com.

Department of Experiment & Teaching, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: wuhuihao99@163.com.

Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: 78926946@qq.com.

Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: 87032164@qq.com.

Engineering Research Center of Key Technology and Industrialization of Cell-based Vaccine, Ministry of Education, Biomedical Research Center, Northwest Minzu University, Lanzhou, 730030, China; Gansu Tech Innovation Center of Animal Cell, Biomedical Research Center, Northwest Minzu University, 730030, Lanzhou, China. Electronic address: qiaozilin@xbmu.edu.cn.